Crossover Inhibition
Because of rectification at the synapse, the negative parts of the ON and OFF patterns are lost in the excitatory and inhibitory currents. But crossover restores the missing components: What is lost from the ON input is restored by the OFF system, and what is lost from the OFF input is restored by the ON system.

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Cartoon of the retina showing layering of neurons

Directional Selectivity

Feedback and Crossover inhibition

Multiple Representations of the Visual Scene

Regions of space/time frequency

Space-time rasters for ON and OFF cells

Patching a neuron in a retinal slice

Targeting Retinal Neuron Subregions with Arficial Rhodopsins